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ISSN: 0001-5113 

AADRAY 

ACTA ADRIAT., 

51(1): 9 - 33, 2010 

UDC: 582.272 (497.5) (262.3) 

A checklist of the benthic marine macroalgae from the eastern 

Adriatic coast: II. Heterokontophyta: Phaeophyceae 

Boris ANTOLIĆ*, Ante ŠPAN, Ante ŽULJEVIĆ, Vedran NIKOLIĆ, 

Ivana GRUBELIĆ, Marija DESPALATOVIĆ and Ivan CVITKOVIĆ 

Institute of Oceanography and Fisheries, Laboratory for Benthos 

Šet. I. Meštrovića 63, 21 000 Split, Croatia 

*Corresponding author, e-mail: antolic@izor.hr 

A checklist of brown algae (Heterokontophyta: Phaeophyceae) from the eastern Adriatic coast, 

based on records published from 1948 to 1997, is presented. For geographic analysis the coast is 

divided into three parts: northern, middle, and southern. A total of 180 macroalgal taxa at specific 

and infraspecific levels are recognized, in which 12 new combinations are proposed. 

Key words: marine benthic macroalgae, Heterokontophyta, Phaeophyceae, checklist, eastern Adri- 

atic coast 

INTRoDUCTIoN 

The present checklist of brown algae from 

the eastern Adriatic coast is intended to be a 

part of the catalogue of benthic algal taxa of the 

eastern Adriatic coast which includes systematic 

divisions Rhodophyta, Heterokontophyta and 

Chlorophyta. The Chlorophyta were treated in 

the first part (ANTOLIĆ et al., 2001). A first checklist 

of Adriatic macroalgae was compiled by 

GIACCONE (1978). That list, however, has no data 

for the southern Adriatic. A more recent check 

list (RIBERA et al., 1992), which covers brown 

algae of the entire Mediterranean Sea, treats the 

Adriatic as a single region and some taxa are 

omitted. Most recently, FURNARI et al. (1999) published 

a catalogue of the benthic marine macroalgae 

of the western Adriatic coast, providing 

more detailed information for that region than 

may be obtained from existing checklists. 

Our present checklist has been compiled 

following the scheme used in the first part of 

this series (ANTOLIĆ et al., 2001). For geographic 

analysis, the eastern Adriatic coast has been 

divided into three parts (Fig. 1). The northern 

part (NEAd) extends from the Gulf of Trieste in 

Italy, along the coast of Slovenia to Jablanac in 

Croatia, the middle part (MEAd) from Jablanac 

to Gradac in Croatia, and the southern part 

(SEAd) from Gradac, along the coast of Montenegro 

to Vlorë in Albania. 

The following references were used in preparing 

this checklist: 1 - ANTOLIĆ et al. (1995); 

2 - AVČIN et al. (1974); 3 - ERCEGOVIĆ (1948); 

4 - ERCEGOVIĆ (1952); 5 - ERCEGOVIĆ (1955a); 

6 - ERCEGOVIĆ (1955b); 7 - ERCEGOVIĆ (1957); 

8 - ERCEGOVIĆ (1966); 9 - ERCEGOVIĆ (1968); 

10 - FLETCHER et al. (1988); 11 - GIACCONE (1978); 

12 - MATJAŠIĆ et al. (1975); 13 - MUNDA (1954); 

14 - MUNDA (1960); 15 - MUNDA (1979); 16 - PIG- 

NATTI & GIACCONE (1967); 17 - SOLAZZI (1971); 

18 - SILVA et al. (1996); 19 - ŠERMAN et al. (1981); 

20 - ŠPAN (1972); 21 - ŠPAN (1980); 22 - ŠPAN & 

ANTOLIĆ (1981); 23 - ŠPAN & ANTOLIĆ (1983); 24

10 ACTA ADRIATICA, 51(1): 9 - 34, 2010 

- ŠPAN & ANTOLIĆ (1988); 25 - ŠPAN & ANTOLIĆ 

(1994); 26 - ŠPAN & ANTOLIĆ (1997); 27 - ŠPAN 

et al. (1996); 28 - VATOVA (1948); 29 - VUKOVIČ 

(1980); 30 - ZAVODNIK et al. (1981). 

Fig.1. Map of the investigated area 

For each part of the eastern Adriatic coast, 

records are shown by numbers that correspond 

to bibliographic references published from 1948 

to 1997 (Table 1). 

Italics are used for accepted algal taxa, with 

roman type for synonyms, misapplied names 

and doubtful or unaccepted taxa. Superscript 

numbers in the brackets refer to the Notes. 

Authors of names are given in full. Authors of 

synonyms are reported as quoted in original 

papers. 

The taxonomy is arranged according to the 

website www.algaebase.org (GUIRY & GUIRY, 

2009). 

It is important to emphasize that we have 

included in this checklist 12 taxa from genera 

Cystoseira, Ectocarpus and Elachista, designated 

by ERCEGOVIĆ (1948, 1952, 1955a, 1955b, 

1957) as new for science, in the renamed form. 

These taxa were considered by some scientists 

as taxonomic synonyms of existing taxa or 

did not register in their inventories (CORMACI 

et al., 1992; RIBERA et al., 1992; GUIRY & GUIRY, 

2009). We are doing so because we believe that 

renaming should be based on both genetic and 

very detailed comparative studies of new algal 

material. 

Moreover, in this list we include the taxa 

of genera Padinopsis, Adriogloia and Dalmatogloia 

which are in some checklists included as 

taxa inquirenda (RIBERA et al., 1992). 

An alphabetical list of taxa (Annex) and taxa 

inquirenda are given. 

A total of 180 benthic macroalgal taxa at 

specific and infraspecific levels are included in 

the list (Table 2). 

In comparison to the 160 taxa recorded in 

the entire Adriatic Sea by RIBERA et al. (1992), and 

140 taxa recorded for the western Adriatic coast 

by FURNARI et al. (1999), we recorded about 11% 

and 22% more taxa, respectively, although we 

introduced some taxa that these authors listed 

as taxonomic synonyms, or they did not mention, 

or that they listed in the category of taxa 

inquirenda. 

In our study, a higher number of algal taxa 

were recorded in the middle (153) and in the 

northern (129) parts of the Adriatic, while only 

75 taxa were recorded in the southern part. 

Table 1. Numbers of references used for the three parts of the eastern Adriatic coast: northern (NEAd), middle (MEAd), 

and southern (SEAd) 

Parts of Adriatic Reference numbers 

NEAd 2 4 10 11 12 13 14 15 16 18 20 27 28 29 30 

MEAd 3 4 5 6 7 8 9 11 18 20 21 25 26 

SEAd 1 3 4 6 17 19 20 22 23 24

AntOlIć et al.: Checklist of macroalgae from the eastern Adriatic coast: II. Heterokontophyta: Phaeophyceae 

Table 2. Taxonomic list of benthic marine macroalgal taxa from the three parts of the eastern Adriatic coast (northern - 

NEAd, middle - MEAd, southern - SEAd). For the meaning of numbers see the text 

Taxa NEAd MEAd SEAd 

HETEROKONTOPHYTA 

PHAEOPHYCEAE Kjellman (FUCOPHYCEAE Warming) 

C u t l e r i a l e s Bessey 

Cutleriaceae Griffiths et Henfrey 

Cutleria Greville 

C. adspersa (Mertens ex Roth) De Notaris (1) ………………………....... 11 8 - 

C. chillosa (Falkenberg) P.C.Silva (2) ....................................................... 

= Cutleria monoica Ollivier 

11 6 5 

C. multifida (Turner) Greville (3) ............................................................... 28 8 23 

Zanardinia Zanardini 

Zanardinia typus (Nardo) P.C.Silva …………………………………..... 

= Zanardinia prototypus (Nardo) Nardo 

= Zanardinia collaris (C.Agardh) P.L. Crouan et 

H.M. Crouan 

D e s m a r e s t i a l e s Setchell et N.L.Gardner 

Arthrocladiaceae Chauvin 

28 7 17 

Arthrocladia Duby 

A. villosa (Hudson) Duby ......................................................................... 11 7 22 

Desmarestiaceae (Thuret) Kjellmann 

Desmarestia J.V.Lamouroux 

D. ligulata (Stackhouse) J.V. Lamouroux ................................................ 

= Desmarestia adriatica Ercegović 

D i c t y o t a l e s Bory de Saint-Vincent 

Dictyotaceae J.V.Lamouroux ex Dumortier 

Dictyopteris J.V.Lamouroux 

D. polypodioides (A.P.De Candolle) J.V.Lamouroux .……………….... 

= Haliseris polypodioides (A.P. de Candolle) C.Agardh 

= Dictyopteris membranacea (Stackhouse) Batters 

- 3 - 

28 7 19 

Dictyota J.V.Lamouroux 

D. dichotoma (Hudson) J.V.Lamouroux 

var. dichotoma ………………………………………………......... 28 7 17 

var. intricata (C.Agardh) Greville ……………………………....... 

= Dictyota implexa (Desfontaines) J.V.Lamouroux 

= Dictyota dichotoma var. implexa (Desfontaines) 

S.F.Gray 

= Dictyota dichotoma f. implexa (Desfontaines) Hauck 

13 8 19 

11


Table 2. cont’d 

Taxa 

D. fasciola (Roth) J.V.Lamouroux 

NEAd MEAd SEAd 

var. fasciola ………………………………………………….......... 28 7 22 

= Dilophus fasciola (Roth) M.Howe 

var. repens (J.Agardh) Ardissone ……………………………....... 

= Dilophus repens (J.Agardh) J.Agardh 

= Dilophus fasciola var. repens (J.Agardh) J.Feldmann 

- 11 - 

D. linearis (C.Agardh) Greville ……………………………………....... 14 7 22 

D. mediterranea (Schiffner) G.Furnari .................................................... - 11 - 

= Dilophus mediterraneus Schiffner 

D. spiralis Montagne ………………………………………………...... 

= Dilophus ligulatus (Kützing) J.Feldmann 

= Dilophus spiralis (Montagne) G.Hamel 

Padina Adanson 

P. pavonica (Linnaeus) Thivy ………………………………………..... 

= Padina pavonia J.V.Lamouroux 

16 8 17 

28 7 17 

Padinopsis Ercegović 

P. adriatica Ercegović (4) …………………………………………......... - 6 - 

Spatoglossum Kützing 

S. solieri (Chauvin ex Montagne) Kützing …………………………...... - 26 - 

Taonia J.Agardh 

T. atomaria (Woodward) J.Agardh …………………………………..... 28 7 23 

D i s c o s p o r a n g i a l e s Schmidt 

Choristocarpaceae Kjellmann 

Choristocarpus Zanardini 

C. tenellus Zanardini ……………………………………....................... 28 6 - 

Discosporangium Falkenberg 

D. mesarthrocarpum (Meneghini) Hauck …………………………....... - 7 - 

E c t o c a r p a l e s Setchell et N.L.Gardner 

Acinetosporaceae G.Hamel ex J. Feldmann 

Acinetospora Bornet 

A.crinita (Carmichael) Kornmann ........................................................... 

= Ectocarpus crinitus Carmichael 

= Acinetospora vidovichii (Meneghini) Sauvageau 

= Acinetospora pusilla (Griffiths ex Harvey) De Toni 

Feldmannia G.Hamel 

F. battersiides (Ercegović) Cormaci et G.Furnari 

f. battersiides ………………………………………………....... 

= Ectocarpus battersiides Ercegović 

f. maior (Ercegović) Antolić et Špan, comb. nov. ………........... 

= Ectocarpus battersiides Ercegović f. maior Ercegović 

14 

7 

23 

- 5 - 

- 5 -




f. spongiosessilis (Ercegović) Antolić et Špan, comb. nov. ......... 

= Ectocarpus battersiides Ercegović f. spongiosessilis 

Ercegović 

27 5 - 

f. taoniae (Ercegović) Antolić et Špan, comb. nov. ......……...... 

= Ectocarpus battersiides Ercegović f. taoniae Ercegović 

F. irregularis (Kützing) G.Hamel 

- 5 - 

var. irregularis ………………………………………………......... 

= Ectocarpus irregularis Kützing 

= Giffordia irregularis (Kützing) Joly 

28 5 22 

var. lebeliides (Ercegović) Antolić et Špan, comb. nov..................... 

= Ectocarpus irregularis subsp. lebeliides Ercegović 

27 5 - 

F. lebelii (J.E. Areschoug ex P.L. Crouan et H.M. Crouan) G. Hamel..... 

= Feldmannia caespitula (J.Agardh) Knoepffler-Péguy 

= Ectocarpus lebelii J.E.Areschoug ex P.L.Crouan et 

H.M.Crouan 

= Feldmannia caespitula var. lebelii 

(J.E.Areschoug ex P.L.Crouan et H.M.Crouan) 

Knoepffler-Péguy 

= Ectocarpus paradoxus subsp. lebeliides Ercegović 

16 5 23 

F. padinae (Buffham) G.Hamel ……………………………………....... 

F. paradoxa (Montagne) G.Hamel 

16 - - 

var. paradoxa ………………………………………………........... 

= Ectocarpus paradoxus Montagne 

28 5 22 

f. profonda (Ercegović) Antolić et Špan, comb. nov. ….......... 

= Ectocarpus paradoxus f. profonda Ercegović 

- 7 - 

var. donatiae (Ercegović) Antolić et Špan, comb. nov. …............. 

= Ectocarpus paradoxus var. donatiae Ercegović 

- 7 - 

F. paradoxoides (Ercegović) Cormaci et G. Furnari .………………....... 

= Ectocarpus paradoxoides Ercegović 

- 5 - 

Hincksia J.E.Gray 

H. dalmatica (Ercegović) Cormaci et G.Furnari 

f. dalmatica …………………………………………………..... 

= Ectocarpus dalmaticus Ercegović 

= Giffordia dalmatica (Ercegović) Cormaci et G.Furnari 

f. acinetiformis (Ercegović) Antolić et Špan, comb.nov. ............. 

= Ectocarpus dalmaticus Ercegović f. acinetiformis Ercegović 

H. geniculata (Ercegović) Cormaci et G.Furnari …………………….... 

= Ectocarpus geniculatus Ercegović 

= Giffordia geniculata (Ercegović) Cormaci et G.Furnari 

H. granulosa (J.E.Smith) P.C.Silva ...………………………………....... 

= Ectocarpus granulosus (J.E.Smith) C.Agardh 

= Giffordia granulosa (J.E.Smith) G.Hamel 

H. hauckii (Ercegović) Cormaci et G.Furnari ………………………...... 

= Ectocarpus hauckii Ercegović 

= Giffordia hauckii (Ercegović) Cormaci et G.Furnari 

H. hincksiae (Harvey) P.C.Silva ……………………………………...... 

= Ectocarpus hincksiae Harvey 

= Giffordia hincksiae (Harvey) G.Hamel 

H. mitchelliae (Harvey) P.C.Silva .…………………………………....... 

= Ectocarpus mitchelliae Harvey 

= Giffordia mitchelliae (Harvey) G.Hamel 

2 5 

22 

- 5 - 

- 5 - 

11 5 - 

11 5 - 

15 - - 

16 11 - 

13




H. sandriana (Zanardini) P.C.Silva …………………………………..... 

= Ectocarpus sandrianus Zanardini 

= Giffordia sandriana (Zanardini) G.Hamel 

28 5 - 

Pylaiella Bory 

P. littoralis (Linnaeus) Kjellmann …………………………………........ 16 11 - 

Chordariaceae Greville 

Acrospongium Schiffner 

A. ralfsioides Schiffner ……………………………………………….... 28 11 - 

Adriogloia Ercegović 

A. adriatica Ercegović (5) …………………………………………........ - 6 - 

Ascocyclus Magnus 

A.orbicularis (J. Agardh) Kjellman …………………………………..... 

= Myrionema orbiculare J. Agardh 

= Myrionema magnusii (Sauvageau) Loiseaux 

= Ascocyclus magnusii Sauvageau 

Asperococcus J.V.Lamouroux 

A. bullosus J.V.Lamouroux …………………………………………...... 

= Asperococcus turneri (Dillwyn ex J.E.Smith) W.J.Hooker 

28 7 22 

28 7 17 

A. ensiformis (Chiaje) M.J. Wynne …………………………………...... 

= Asperococcus compressus A.W.Griffiths ex W.J.Hooker 

= Haloglossum compressum (A.W.Griffiths ex W.J.Hooker) 

G.Hamel 

28 8 23 

A. fistulosus (Hudson) W.J.Hooker …………………………………..... 

= Asperococcus echinatus (Mertens ex Roth) Greville 

- 7 - 

A. scaber Kuckuck …………………………………………………....... 28 8 - 

Cladosiphon Kützing 

C. contortus (Thuret) Kylin …………………………………………..... 28 - - 

C. cylindricus (Sauvageau) Kylin …………………………………........ 

= Castagnea cylindrica Sauvageau 

11 - - 

C. mediterraneus Kützing …………………………………………........ 

= Castagnea mediterranea (Kützing) Hauck 

11 8 22 

C. zosterae (J.Agardh) Kylin ………………………………………....... 

= Castagnea zosterae (J.Agardh) Thuret 

16 - - 

Corynophlaea Kützing 

C. flaccida (C.Agardh) Kützing ……………………………………....... 

= Myriactis adriatica (J.Agardh) De Toni 

= Leathesia flaccida (C.Agardh) Endlicher 

C.umbellata (C.Agardh) Kützing ……………………………………..... 

= Corynephora umbellata C.Agardh 

= Leathesia umbellata (C.Agardh) Meneghini 

11 - - 

28 8 - 

Cylindrocarpus P.L. Crouan et H.M. Crouan 

C. microscopicus P.L. Crouan et H.M. Crouan ……………………....... 11 - - 

Dalmatogloia Ercegović 

D. bryozoi Ercegović (6) ……………………………………………....... - 6 -




Elachista Duby 

E. fucicola (Velley) J.E.Areschoug 

f. fucicola …………………………………………………........ 27 11 19 

f. profunda Ercegović ………………………………………..... 

E. intermedia P.L. Crouan et H.M. Crouan 

- 7 - 

var. intermedia ……………………………………………............. 28 11 19 

= Elachista kuckuckiana Schiffner 

f. profunda Ercegović ……………………………………......... - 3 - 

var. clavaeformis Ercegović ………………….........…………….. 

E. neglecta Kuckuck nom. illeg. 

- 3 - 

var. neglecta ……………………………………………………..... 28 7 23 

var. jabukae (Ercegović) Antolić et Špan, comb. nov. ..................... 

= Elachista jabukae Ercegović 

= Elachista neglecta subsp. jabukae Ercegović 

- 3 - 

E. stellaris J.E.Areschoug …………………………………………........ 28 - 23 

Giraudia Derbès et Solier 

G. sphacelarioides Derbès et Solier ....................……………………..... 28 7 22 

Hecatonema Sauvageau 

H. terminale (Kützing) Kylin .................................................................... 

= Hecatonema maculans (F.S.Collins) Sauvageau 

Herponema J.Agardh 

H. valiantei (Bornet ex Sauvageu) G.Hamel ………………………....... 

= Ectocarpus valiantei Bornet ex Sauvageau 

Kuetzingiella Kornmann 

K. battersii (Bornet ex Sauvageau) Kornmann 

var. battersii …………………………………………………......... 

= Ectocarpus battersii Bornet ex Sauvageau 

= Feldmannia battersii (Bornet ex Sauvageau) G.Hamel 

var. mediterranea (Bornet ex Sauvageau) Gómez et Ribera …........ 

= Ectocarpus battersii var. mediterraneus Bornet ex Sauvageau 

- 26 - 

- 9 - 

11 5 - 

16 - - 

Leathesia S.F.Gray 

L. mucosa J.Feldmann 

var. mucosa ……………………………………………………...... 11 3 - 

f. exuberans Ercegović ……………………………………....... - 5 - 

Leptonematella P.C.Silva 

L. fasciculata (Reinke) P.C.Silva ……………………………………..... 

= Leptonema fasciculatum Reinke 

Liebmannia J.Agardh 

L. leveillei J.Agardh …………………………………………………..... 

= Mesogloia leveillei Meneghini nom.illeg. 

= Mesogloea leveillei (J.Agardh) Meneghini 

Litosiphon Harvey 

L. laminariae (Lyngbye) Harvey ……………………………………..... 

= Streblonema oligosporum Strömfelt 

= Entonema oligosporum (Strömfelt) Kylin 

28 - - 

28 9 - 

- 5 - 

15




Mesogloia C.Agardh 

M. vermiculata (J.E.Smith) S.F.Gray ..……………………………….... 

= Mesogloea vermiculata (J.E.Smith) Le Jolis nom.illeg. 

28 - 22 

Microspongium Reinke 

M. kuckuckianum Schiffner …………………………………………..... 28 11 - 

Myriactula Kuntze 

M.elongata (Sauvageau) G.Hamel …………………………………...... 

= Myriactis elongata Sauvageau 

M.microscopica (Ercegović) Ercegović ……………………………...... 

= Myriactis microscopica Ercegović 

M. rigida (Sauvageau) G.Hamel ……………………………………..... 

= Myriactis rigida Sauvegau 

M. rivulariae (Suhr) J.Feldmann …………………………………….... 

= Myriactis pulvinata Kützing 

= Myriactula pulvinata (Kützing) Kuntze 

M. stellulata (Harvey) Levring ……………………………………....... 

= Myriactis stelullata (Harvey) Batters 

Myrionema Greville 

M. liechtensternii Hauck ……………………………………………...... 

= Phaeosphaerium liechtensternii (Hauck) De Toni 

M. strangulans Greville ……………………………………………....... 

= Myrionema vulgare Thuret 

27 3 23 

- 3 23 

- 3 23 

28 3 3 

27 7 23 

11 - - 

28 8 23 

Myriotrichia Harvey 

M. adriatica Hauck …………………………………………………...... 

M. clavaeformis Harvey 

11 25 - 

f. clavaeformis ........................................................................... 

= Myriotrichia repens Hauck 

= Streblonema sphaericum (Derbès et Solier) Thuret 

28 7 23 

= Dichosporangium repens (Hauck) Hauck 

f. acycla (Ercegović) Antolić et Špan, comb. nov. ………........ 

= Myriotrichia repens Hauck f. acycla Ercegović 

Nemacystus Derbès et Solier 

N. flexuosus (C.Agardh) Kylin 

var. giraudyi (J.Agardh) Y.S.D.M.De Jong ....……………….... 

= Nemacystus ramulosus Derbès et Solier 

Petrospongium Nägeli 

P. berkeleyi (Greville) Nägeli ex Kützing …………………………....... 

= Cylindrocarpus berkeleyi (Greville) P.L.Crouan et 

H.M.Crouan 

Protasperococcus Sauvageau 

P. myriotrichiiformis Sauvageau ……………………………………..... 

= Myriotrichia protasperococcus Kuckuck 

- 6 - 

28 7 - 

11 - - 

28 7 -




Punctaria Greville 

P. latifolia Greville …………………………………………………..... 28 8 - 

P. tenuissima (C. Agardh) Greville …………………………………..... 

= Streblonema effusum Kylin 

= Entonema effusum (Kylin) Kylin 

- 7 - 

Sauvageaugloia G.Hamel ex Kylin 

S. divaricata (Clemente) Cremades …………………………………..... 

= Sauvageaugloia griffithsiana (Greville ex W.J.Hooker) 

G.Hamel ex Kylin 

= Mesogloia griffithsiana Greville ex W.J.Hooker 

11 21 - 

Spermatochnus Kützing 

S. paradoxus (Roth) Kützing 

var. paradoxus ………………………………………..................... 29 7 19 

var. adriaticus Ercegović …………………………......................... 11 3 3 

Stictyosiphon Kützing 

S. adriaticus Kützing ………………………………………………....... 28 7 24 

S. tortilis (Gobi) Reinke ................……………………………………... 14 - - 

Stilophora J.Agardh 

S. tenella (Esper) P.C.Silva ..................................................................... 

= Stilophora rhizodes (C.Agardh) J.Agardh nom. illeg. 

Streblonema Derbès et Solier in Castagne 

S. infestans (H.Gran) Batters ………………………………………....... 

= Endodictyon infestans H.Gran 

28 7 22 

- 7 - 

Streblonemopsis R. Valiante 

S. irritans R. Valiante .............................................................................. - 9 - 

Striaria Greville 

S. attenuata (Greville) Greville ............................................................... 28 9 - 

Zosterocarpus Bornet 

Z. oedogonium (Meneghini) Bornet .......................................................... 11 7 - 

Ectocarpaceae C.Agardh 

Ectocarpus Lyngbye 

E. bombycinus Kützing ……………………………………………........ 

E. fasciculatus Harvey 

- 18 - 

var. fasciculatus ……………………………………….................. 16 - - 

var. abbreviatus (Kützing) Sauvageau …………………............... 28 - - 

= Ectocarpus abbreviatus Kützing 

E. flagelliformis Kützing ……………………………………………...... 

= Ectocarpus fasciculatus Kützing nom. illeg. 

= Ectocarpus approximatus Kützing 

18 18 - 

E. parvulus Kützing …………………………………………………..... 18 - - 

E. pectinis Ercegović ………………………………………………....... - 5 - 

E. rufulus Kützing …………………………………………………........ 18 - - 

17



Taxa 

E. siliculosus (Dillwyn) Lyngbye 

NEAd MEAd SEAd 

var. siliculosus ………………………………………………......... 

= Ectocarpus confervoides (Roth) Le Jolis nom. illeg. 

= Ectocarpus siliculosus var. confervoides (Roth) Kjellman 

28 5 22 

var. adriaticus (Ercegović) Cormaci et G.Furnari (7) …………....... 

= Ectocarpus adriaticus Ercegović 

11 5 22 

f. maior (Ercegović) Antolić et Špan, comb..nov. ...…............... 

= Ectocarpus adriaticus Ercegović f. maior Ercegović 

- 6 - 

var. arctus (Kützing) T.Gallardo (8) …………………………......... 14 7 - 

= Ectocarpus arctus Kützing 

var. dasycarpus (Kuckuck) T.Gallardo ………………………......... 

= Ectocarpus dasycarpus Kuckuck 

= Ectocarpus siliculosus var. crassus (Kjellman) T.Gallardo 

var. pygmaeus (J.E.Areschoug) T.Gallardo (9) ………………......... 

= Ectocarpus pygmaeus J.E.Areschoug 

var. subulatus (Kützing) T.Gallardo (10) ………………………........ 

= Ectocarpus subulatus Kützing 

var. venetus (Kützing) T.Gallardo (11) ……………………............ 

= Ectocarpus venetus Kützing 

11 - - 

11 - - 

11 - - 

11 11 - 

E. simpliciusculus C.Agardh 

f. simpliciusculus ………………………………………………. 18 5 - 

f. reductus (“reducta”) Ercegović ……………………………... - 7 - 

E. virescens Thuret ex Sauvageau …………………………………....... - 5 - 

Kuckuckia G.Hamel 

K. spinosa (Kützing) Kornman ……………………………………......... 

= Ectocarpus spinosus Kützing 

F u c a l e s Kylin 

Cystoseiraceae Kützing 

28 25 - 

Cystoseira C.Agardh 

C.amentacea (C.Agardh) Bory 

var. spicata (Ercegović) Giaccone ……………………………........ 

= Cystoseira spicata Ercegović 

= Cystoseira stricta Sauvageau var. spicata (Ercegović) 

Giaccone 

C. barbata (Stackhouse) C.Agardh 

4 4 4 

var. barbata ………………………………………………….......... 

= Cytoseira barbata J.Agardh 

= Cystoseira barbata (Goodenough et Woodward) C.Agardh 

28 4 4 

f. insularum Ercegović ……………………………………........ - 4 - 

var. tophuloidea (Ercegović) Giaccone ............................................ 

= Cystoseira barbata subsp. tophuloidea Ercegović 

= Cystoseira barbata var. tophuloidea (Ercegović) Giaccone 

C. compressa (Esper) Gerlof et Nizamudin 

11 4 - 

f. compressa ………………………………................................. 

= Cystoseira fimbriata (Desfontaines) Bory 

= Cystoseira abrotanifolia C.Agardh 

28 4 4 

f. plana (Ercegović) Cormaci et al. .............................................. 

= Cystoseira abrotanifolia subsp. plana Ercegović 

15 4 4




f. rosetta (Ercegović) Cormaci et al. ............................................ 

= Cystoseira abrotanifolia subsp. rosetta Ercegović 

= Cystoseira fimbriata (Desfontaines) Bory f. rosetta 

4 4 4 

f. insularum (Ercegović) Antolić et Špan, comb. nov. …............. 

= Cystoseira abrotanifolia C.Agardh f. insularum Ercegović 

C. corniculata (Turner) Zanardini 

- 4 - 

var. corniculata ……………………………………………............. 

= Cystoseira corniculata Hauck 

28 4 4 

f. imperfecta Ercegović (12) …………………………………....... - 4 - 

var. divergens (Ercegović) Antolić et Špan, comb. nov. ................. 

= Cystoseira corniculata subsp. divergens Ercegović 

- 3 - 

var. laxior (Ercegović) Antolić et Špan, comb. nov. ........................ 

= Cystoseira corniculata subsp. laxior Ercegović 

C. crinita Duby 

30 4 4 

f. crinita ……………………………………………………....... 

= Cystoseira crinita Bory 

28 4 4 

f. semispinosa Ercegović ……………………………………..... - 4 4 

C. crinitophylla Ercegović …………………………………………....... 12 4 22 

C. dubia R.Valiante …………………………………………………...... 

= Cystoseira fucoides Ercegović 

C. foeniculacea (Linnaeus) Greville 

11 4 - 

f. foeniculacea ………………………………………………...... 

= Fucus discors Linnaeus 

= Cystoseira discors (Linnaeus) C.Agardh 

= Cystoseira ercegovicii Giaccone 

f. latiramosa (Ercegović) Gómez, Garreta, 

28 4 4 

Barceló, Ribera et Rull Lluch ............ 

= Cystoseira discors subsp. latiramosa Ercegović 

= Cystoseira discors f. latiramosa (Ercegović) Giaccone 

= Cystoseira ecegovicii Giaccone f. latiramosa (Ercegović) 

Giaccone 

f. tenuiramosa (Ercegović) Gómez, Garreta, 

4 4 4 

Barceló, Ribera et Rull Lluch …......... 

= Cystoseira discors f. tenuiramosa Ercegović 

= Cystoseira ercegovicii Giaccone f. tenuiramosa 

C. humilis Kützing 

16 4 - 

var. humilis …………………………………………………........... 

= Cystoseira abrotanifolia subsp. pustulata Ercegović 

= Cystoseira compressa var. pustulata Ercegović 

4 4 4 

var. myriophylloides (Sauvageau) J.H.Price et D.M. John …........... 

= Cystoseira myriophylloides Sauvageau 

C. jabukae Ercegović 

11 - - 

f. jabukae ….................................................................................. 

f. tenuissima (Ercegović) Cormaci, G.Furnari, Giaccone, 

- 4 - 

Scammacca et Serio .......... 

= Cystoseira jabukae subsp. tenuissima Ercegović 

- 4 - 

C. pelagosae Ercegović ............................................................................ - 4 - 

C. sauvageauana G.Hamel ...................................................................... 

= Cystoseira selagenoides R.Valiante 

C.spinosa Sauvageau 

- - 17 

var. spinosa ....…………………………………...…....................... 

= Cystoseira erica-marina R.Valiante 

= Cystoseira adriatica Sauvageau 

28 9 4 

19




var. compressa (Ercegović) Cormaci et al. ...................................... 

= Cystoseira adratica subsp. compressa Ercegović 

= Cystoseira adriatica var. compressa (Ercegović) 

Giaccone in Amico et al. 

= Cystoseira adriatica subsp. intermedia Ercegović 

= Cystoseira adriatica var. intermedia (Ercegović) 


= Cystoseira platyramosa Ercegović 

11 4 4 

var. tenuior (Ercegović) Cormaci et al. ........................................... 

= Cystoseira adriatica subsp. tenuior Ercegović 

= Cystoseira adriatica f. tenuior (Ercegović) 


= Cystoseira adriatica subsp. reducta Ercegović 

= Cystoseira adriatica f. reducta (Ercegović) 


11 4 4 

C. squarrosa De Notaris ……………………………………………..... 

= Cystoseira spinosa var. squarrosa (De Notaris) Giaccone 

11 8 4 

C. zosteroides C.Agardh ……………………………………………...... 

= Cystoseira opuntioides Bory ex Montagne 

- 4 - 

Fucaceae Adanson 

Fucus Linnaeus 

F. virsoides J.Agardh ………………………………………………........ 28 8 17 

Sargassaceae Kützing 

Sargassum C.Agardh 

S. acinarium (Linnaeus) Setchell (13) ……………...………………….... 

= Sargassum vulgare var. linifolium C.Agardh 

= Sargassum linifolium C.Agardh 

28 8 - 

S. hornschuchii C.Agardh ................................………………………..... 28 7 20 

S. vulgare C.Agardh 

var. . vulgare …………………………………………………........ 

= Sargassum salicifolium Naccari, 

= Sargassum vulgare var. salicifolium C.Agardh 

= Sargassum vulgare subsp. megalophyllum (Montagne) Grunow 

L a m i n a r i a l e s Kylin 

Laminariaceae Bory 

20 7 17 

Laminaria J.V.Lamouroux 

L. rodriguezii Bornet ………………………………………………......... - 7 - 

R a l f s i a l e s Nakamura 

Ralfsiaceae Farlow 

Pseudolithoderma Svedelius 

P. adriaticum (Hauck) Verlaque ……………………………………....... 

= Lithoderma adriaticum Hauck 

16 7 1




Ralfsia Berkeley in J.E.Smith et Sowerby 

R. verrucosa (J.E.Areschoug) J.E.Areschoug………………………....... 16 7 19 

S c y t o s i p h o n a l e s J.Feldmann 

Scytosiphoniaceae Farlow 

Colpomenia (Endlicher) Derbès et Solier 

C. sinuosa (Mertens ex Roth) Derbès et Solier ......................................... 28 8 19 

Compsonema Kuckuck 

C. gracile Kuckuck ................................................................................... 28 11 - 

C. saxicolum (Kuckuck) Kuckuck ............................................................ 

= Myrionema saxicola Kuckuck 

10 - - 

Hydroclathrus Bory de Saint-Vincent 

H. clathratus (C.Agardh) M.A.Howe ………………………………....... - 11 - 

Petalonia Derbès et Solier 

P. fascia (O.F.Müller) Kuntze .................................................................. 

= Phyllitis fascia (O.F.Müller) Kützing 

P. zosterifolia (Reinke) Kuntze ……………………………………….... 

= Phyllitis zosterifolia Reinke 

28 8 - 

28 - - 

Scytosiphon C.Agardh 

S. dotyi M.J.Wynne …………………………………………………...... 11 - - 

S. lomentaria (Lyngbye) Link (14) ………...…………………………...... 

= Scytosiphon lomentarius (Lyngbye) J.Agardh 

= Scytosiphon simplicissimus (Clemente) Cremades 

28 8 21 

S p h a c e l a r i a l e s Oltmanns 

Sphacelariaceae Decaisne 

Cladostephus C.Agardh 

C. spongiosum (Hudson) C.Agardh 

f. verticillatum (Lightfoot) Prud’homme van Reine .................... 

= Cladostephus verticillatus (Lightfoot) Lyngbye 

= Cladostephus hirsutus (Linnaeus) C.F.Boudouresque 

et M.Perret-Boudouresque 

Sphacella Reinke 

S. subtilissima Reinke ………………………………………………....... - 9 - 

Sphacelaria Lyngbye in Hornemann 

S. cirrosa (Roth) C.Agardh …………………………………………....... 

= Sphacellaria hystrix Suhr ex Reinke 

= Sphacelaria irregularis Kützing 

= Sphacelaria pennata Lyngbe 

= Sphacelaria cirrosa var. pennata (Lyngbe) Hauck 

28 7 22 

S. fusca (Hudson) S.F.Gray …………………………………………...... 11 7 22 

28 

7 

22 

21




S. nana Nägeli ex Kützing ……………………………………………..... 

= Sphacelaria britannica Sauvageau 

16 - - 

S. plumula Zanardini ................................................................................. 28 7 19 

S. rigidula Kützing …………………………………………………........ 

= Sphacelaria furcigera Kützing 

16 25 22 

S. tribuloides Meneghini ……………………………………………....... 28 7 19 

Stypocaulaceae Oltmans 

Halopteris Kützing 

H. filicina (Grateloup) Kützing ………………………………………..... 

= Sphacelaria filicina (Grateloup) C.Agardh 

Stypocaulon Kützing 

Stypocaulon scoparium (Linnaeus) Kützing ………………………....... 

= Halopteris scoparia (Linnaeus) Sauvageau 

= Sphacelaria scoparia (Linnaeus) Lyngbye 

S p o r o c h n a l e s Sauvageau 

Sporochnaceae Greville 

Carpomitra Kützing 

C. costata (Stackhouse) Batters 

var. costata …………………………………………………............ 

= Fucus costatus Stackhouse 

= Carpomitra cabrerae (Clemente) Kützing 

var. dichotoma (Zanardini) J.Feldmann ............................................ 

= Sporochnus dichotomus Zanardini 

28 7 22 

13 7 17 

- 9 - 

- 7 - 

Nereia Zanardini 

N. filiformis (J.Agardh) Zanardini …………………………………….... 28 7 22 

Sporochnus C.Agardh 

S. pedunculatus (Hudson) C.Agardh ………………………………… 28 7 19


NoTeS 

1. This species includes the sporophytic stage 

Aglaozonia melanoidea Sauvageau nom.inval. 


Aglaozonia chilosa Falkenberg. 


Aglaozonia parvula (Greville) Zanardini. 

4. RIBERA et al. (1992) and GUIRY & GUIRY (2009) 

considered this species as taxon inquirendum, but 

we will accept it and include it in this checklist. 

5. We followed GUIRY & GUIRY (2009) including 

this species in the checklist as a currently 

accepted taxon; RIBERA et al. (1992) treated this 

species as taxon inquirendum. 

6. We followed GUIRY & GUIRY (2009) including 

this species in the checklist as a currently 

accepted taxon; RIBERA et al. (1992) treated this 

species as taxon inquirendum. 

7. GIACCONE (1978) transferred the species 

Ectocarpus adriaticus Ercegović into the variety 

Ectocarpus siliculosus (Dillwyn) Lyngbye 

var. adriaticus (Ercegović) Giaccone without 

specifying a bibliographic reference of the 

basionym. This is incorrect according to Art. 

33.2 of the International Code of Botanical 

Nomenclature (McNEILL et al., 2006). The 

new combination was correctly proposed by 

CORMACI & FURNARI (1987) and we included it 

in this checklist 


Ectocarpus arctus Kützing into the variety 

Ectocarpus siliculosus (Dillwyn) Lyngbye var. 

arctus (Kützing) Giaccone without specifying 

a bibliographic reference of the basionym. 

This is incorrect according to Art. 33.2 of the 

International Code of Botanical Nomenclature 

(McNEILL et al., 2006). The new combination was 

correctly proposed by GALLARDO (1992). We 

included this variety in the checklist. GUIRY & 

GUIRY (2009) included it as one of the taxonomic 

synonyms of Ectocarpus siliculosus (Dillwyn) 

Lyngbye. 

23 


Ectocarpus pygmaeus J.E. Areschoug in Kjellman 

into the variety Ectocarpus siliculosus (Dillwyn) 

Lyngbye var. pygmaeus (J.E. Areschoug) 

Giaccone without specifying a bibliographic 

reference of the basionym. This is incorrect 

according to Art. 33.2 of the International Code 

of Botanical Nomenclature (McNEILL et al., 2006). 

The new combination was correctly proposed by 

GALLARDO (1992) and we included this variety 

in the checklist. 


Ectocarpus subulatus Kützing into the variety 


subulatus (Kützing) Giaccone without specifying 

a bibliographic reference of the basionym. The 


GALLARDO (1992). 


Ectocarpus venetus Kützing into the variety 


venetus (Kützing) Giaccone without specifying 

a bibliographic reference of the basionym. The 


GALLARDO (1992). 

12. CORMACI et al. (1992) and RIBERA et al. (1992) 

indicated the form Cystoseira corniculata 

(Turner) Zanardini f. imperfecta Ercegović as 

one of the taxonomic synonyms of the species 

Cystoseira corniculata (Turner) Zanardini. 

GUIRY & GUIRY (2009) included this form by 

the name of Cystoseira corniculata f. imperfecta 

Ercegović in the category “P” (indicates a 

preliminary AlgaeBase entry that has not been 

subjected to any kind of verification). Until 

genetic and detailed comparative research will 

be done on fresh algal material, we consider this 

taxon as distinct. 

13. This species includes the subspecies Sargassum 

salicifolium subsp. linifolium Špan, to which the 

latter was transferred as a variety (ŠPAN, 2005). 

14. Acording to PARENTE et al. (2003) this species 

includes the sporophytic stage Microspongium 

gelatinosum Reinke.


NomeNClUTARAl CHANgeS 

CORMACI & FURNARI (1987) transferred the species 

Ectocarpus battersiides Ercegović into the 

species Feldmannia battersiides (Ercegović) 

Cormaci et G. Furnari. However, ERCEGOVIĆ 

(1955a) described three different forms within 

this species and we suggest new combinations 

as follows: 

Feldmannia battersiides (Ercegović) Cormaci et 

G. Furnari f. maior (Ercegović) Antolić et Špan, 

comb. nov. 

Basionym: Ectocarpus battersiides Ercegović f. 

maior Ercegović in Acta Adriat., 7 (5): 36-38, 

40, Fig. 16. 1955a. 

Feldmannia battersiides (Ercegović) Cormaci 

et G. Furnari f. sporangiosessilis (Ercegović) 

Antolić et Špan, comb. nov. 

Basionym: Ectocarpus battersiides Ercegović 

f. sporangiosessilis Ercegović in Acta Adriat., 7 

(5): 38, 40, Fig. 17 c. 1955a. 

Feldmannia battersiides (Ercegović) Cormaci 

et G. Furnari f. taoniae (Ercegović) Antolić et 

Špan, comb. nov. 

Basionym: Ectocarpus battersiides Ercegović f. 

taoniae Ercegović in Acta Adriat., 7 (5): 38, 40, 

Fig. 17 a, b. 1955a. 

HAMEL (1939) transferred the species Ectocarpus 

irregularis Kützing into the species Feldmannia 

irregularis (Kützing) Hamel. However, 

within this species ERCEGOVIĆ (1955a) described 

the subspecies Ectocarpus irregularis Kützing 

subsp. lebeliides Ercegović, and we suggest a 

new combination as follows: 

Feldmannia irregularis (Kützing) G. Hamel var. 

lebeliides (Ercegović) Antolić et Špan, comb. et 

stat. nov. 

Basionym: Ectocarpus irregularis Kützing 

subsp. lebeliides Ercegović in Acta Adriat., 7 

(5): 54-57, Fig. 25. 1955a. 

HAMEL (1939) transferred the species Ectocarpus 

paradoxus Montagne in Moris et De Notaris 

into the species Feldmannia paradoxa (Montagne) 

G. Hamel. However, ERCEGOVIĆ (1957) 

within the species Ectocarpus paradoxus Montagne 

distinguished the form Ectocarpus para- 

doxus Montagne f. profunda Ercegović and the 

variety Ectocarpus paradoxus Montagne var. 

donatiae Ercegović and we suggest new combinations 

as follows: 

Feldmannia paradoxa (Montagne) G. Hamel 

f. profonda (Ercegović) Antolić et Špan, comb. 

nov. 

Basionym: Ectocarpus paradoxus Montagne 

f. profonda Ercegović in Acta Adriat., 8 (8): 

37-38, Fig. 12 b. 1957. 

Feldmannia paradoxa (Montagne) G. Hamel 

var. donatiae (Ercegović) Antolić et Špan, comb. 

nov. 

Basionym: Ectocarpus paradoxus Montagne 

var. donatiae Ercegović in Acta Adriat., 8 (8): 

38-39, Fig. 12 a. 1957. 

CORMACI & FURNARI (1987) transferred the species 

Ectocarpus dalmaticus Ercegović firstly 

into the species Giffordia dalmatica (Ercegović) 

Cormaci et G. Furnari and later into the species 

Hincksia dalmatica (Ercegović) Cormaci et 

G. Furnari (in GALLARDO, 1992) which is valid 

today. GUIRY & GUIRY (2009) included the species 

Ectocarpus dalmaticus Ercegović in the category 

“U” (indicates a name of uncertain taxonomic 

status, but which has been subjected to 

some verification nomenclaturally). In accepting 

correct renaming of the species by CORMACI & 

FURNARI (1987), we suggest the new combination 

of the form Ectocarpus dalmaticus Ercegović f. 

acinetiformis Ercegović as follows: 

Hincksia dalmatica (Ercegović) Cormaci et G. 

Furnari f. acinetiformis (Ercegović) Antolić et 

Špan, comb. nov. 

Basionym: Ectocarpus dalmaticus Ercegović f. 

acinetiformis Ercegović in Acta Adriat., 7 (5): 

60-62, Fig. 27. 1955a. 

ERCEGOVIĆ (1948) described the new species 

Elachista Jabukae Ercegović. Later, he transfered 

this species into the subspecies Elachista 

neglecta Kuckuck subsp. Jabukae Ercegović 

(ERCEGOVIĆ, 1957). We suggest the new combination 

of Elachista Jabukae Ercegović as 

follows: 

Elachista neglecta Kuckuck var. jabukae 

(Ercegović) Antolić et Špan, comb. et stat. nov.


Basionym: Elachista Jabukae Ercegović in Acta 

Adriat., 3 (5): 10-13, Figs. 4, 5. 1948. 

Acording to PEDERSEN (1978), RIBERA et al. (1992) 

treated the species Myriotrichia repens Hauck as 

one of the taxonomic synonyms of Myriotrichia 

clavaeformis Harvey. However, ERCEGOVIĆ 

(1955b) within the species Myriotrichia repens 

Hauck described six different forms. For one 

form, we suggest a new combination as follows: 

Myriotrichia clavaeformis Harvey f. acycla 

(Ercegović) Antolić et Špan, comb. nov. 

Basionym: Myriotrichia repens Hauck f. acycla 

Ercegović in Acta Adriat., 7 (6): 23-24, Fig. 5d. 

1955b. 

GIACCONE (1978) transferred the species Ectocarpus 

adriaticus Ercegović into the variety 


adriaticus (Ercegović) Giaccone without specifying 

a bibliographic reference of the basionym. 

This is incorrect according to Art. 33.2 of the 


(McNEILL et al., 2006). The new combination was 

correctly proposed by CORMACI & FURNARI 

(1987). In accepting correct renaming of the species 

by CORMACI & FURNARI (1987), we suggest 

the new combination the Ectocarpus adriaticus 

Ercegovic f. maior Ercegovic as follows: 


adriaticus (Ercegović) Cormaci et G.Furnari 

f. maior (Ercegović) Antolić et Špan, comb. et 

stat. nov. 

Basionym: Ectocarpus adriaticus Ercegović f. 

maior Ercegović in Acta Adriat., 7 (5): 16-17. 

1955a. 

GERLOF & NIZAMUDDIN (1975) transferred the 

species Cystoseira abrotanifolia C.Agardh into 

the species Cystoseira compressa (Esper) Gerlof 

et Nizamudin. Until genetic and detailed comparative 

research will be done on new material, 

we suggest the new combination of Cystoseira 

abrotanifolia f. insularum Ercegović, which 

inhabits depths between 1 and 30 m along the 

outer islands of the central Adriatic, as follows: 

25 

Cystoseira compressa (Esper) Gerlof et Nizamudin 

f. insularum (Ercegović) Antolić et Špan, 

comb. nov. 

Basionym: Cystoseira abrotanifolia C.Agardh f. 

insularum Ercegović in Fauna et Flora Adriat., 

2: 102. 1952. 

CORMACI et al. (1992) and RIBERA et al. (1992) cited 

both subspecies Cystoseira corniculata Hauck 

subsp. divergens Ercegović and Cystoseira corniculata 

Hauck subsp. laxior Ercegović Hauck 

as the taxonomic synonyms of Cystoseira corniculata 

(Wulfen) Zanardini. Until genetic and 

detailed comparative research will be done on 

new material, we suggest the new combination 

as follows: 

Cystoseira corniculata (Turner) Zanardini var. 

divergens (Ercegović) Antolić et Špan, comb. et 

stat. nov. 

Basionym: Cystoseira corniculata Hauck subsp. 

divergens Ercegović in Fauna et Flora Adriat., 2: 

18, 106, Icon: Tab. III. 1952. 

Cystoseira corniculata (Turner) Zanardini var. 

laxior (Ercegović) Antolić et Špan, comb. et 

stat. nov. 

Basionym: Cystoseira corniculata Hauck subsp. 

laxior Ercegović in Fauna et Flora Adriat., 2: 17, 

18, 106, Icon: Tab. II. 1952. 

Taxa inquirenda 

Acinetospora species Ercegović: (MEAd: 

ERCEGOVIĆ, 1957). 


divergens Schiffner: (NEAd: GIACCONE, 1978). 


elongatus Schifner: (NEAd: GIACCONE, 1978). 


megacarpus Schiffner: (NEAd: GIACCONE, 

1978). 

Dictyota dichotoma (Hudson) Lamouroux 

f. proliferans Ercegović: (NEAd: ŠPAN et al., 

1996; MEAd: ERCEGOVIĆ, 1957; SEAd: ŠPAN & 

ANTOLIĆ, 1983). 

Dilophus mediterraneus Schiffner var. crassus 

Schiffner: (MEAd: GIACCONE, 1978). 

Cystoseira barabata (Stackhouse) C. Agardh f. 

punctata Ercegović: (NEAd: ERCEGOVIĆ, 1952; 

MEAd: ERCEGOVIĆ, 1952).


Cystoseira discors (Linnaeus) C. Agardh f. 

dubia Ercegović: (MEAd: ERCEGOVIĆ, 1952). 

Cystoseira spicata Ercegović subsp. crassa 

Ercegović: (NEAd: ERCEGOVIĆ, 1952; MEAd: 

ERCEGOVIĆ, 1952; SEAd: ERCEGOVIĆ, 1952). 

Cystoseira spicata Ercegović subsp. elegans 

Ercegović: (MEAd: ERCEGOVIĆ, 1952; SEAd: 

ERCEGOVIĆ, 1952). 

Myriogloea sciurus (Harvey) P.Kuckuck ex 

F.Oltmanns: (NEAd: PIGNATTI & GIACCONE, 

1967). 

Myriotrichia repens Hauck f. brevicellularisplurilocularis 

Ercegović: (MEAd: ERCEGOVIĆ, 

1955b). 

Myriotrichia repens Hauck f. brevicellularisunilocularis 


1955b). 

Myriotrichia repens Hauck f. internodilaismixtolocularis 


1955b). 

Myriotrichia repens Hauck f. longicellularisunilocularis 


1955b). 

Myriotrichia repens Hauck f. longicellularisscoparia 

Ercegović: (MEAd: ERCEGOVIĆ, 1955b). 

Sargassum vulgare C.Agardh f. ercegovicii 

Špan (MEAd: ŠPAN, 1972). 

Sargassum vulgare C.Agardh subsp. jabukae 

Špan (CEAd: ŠPAN, 1972). 

ReFeReNCeS 

ANTOLIĆ, B., A. ŠPAN & E. DRAGANOVIĆ. 1995. Prilog 

poznavanju bentoske flore otoka Mljeta. 

(Contribution to the knowledge of the benthic 

flora of Mljet Island (southern Adriatic, 

Croatia)). Hrvatsko ekološko društvo, Ekol. 

monogr., 6: 531-542. 

ANTOLIĆ, B., A. ŠPAN, A. ŽULJEVIĆ & A. VUKOVIČ. 

2001. Check list of the benthic marine macroalgae 

on the eastern Adriatic coast: I. 

Chlorophyta. Acta Adriat., 42(2): 43-58. 

AVČIN, A., N. MEITH-AVČIN, A. VUKOVIČ & B. 

VRIŠER. 1974. Primerjava bentoških združb 

Strunjanskega in Koprskoga zaliva z obzirom 

na njihove polucijsko pogojene razlike 

(The comparison of benthic communities of 

Strunjan and Kopar Bay due to their differences 

in pollution). Biol. Vestn. Glas. Slov. 

Biol., 22(2): 171-208. 

CORMACI, M., G. FURNARI, G. GIACCONE, B. SCAM- 

MACCA & D.SERIO. 1992. Observations taxonomiques 

et biogéographiques sur quelques 

espèces du genre Cystoseira C. (Taxonmic 

and biogeografical observations on species 

of genus Cystoseira C. Agardh). Bull. Inst. 

Océanogr., 9 (Spécial): 21-35. 

CORMACI, M. & G. FURNARI. 1987. Nomenclatural 

notes on some Mediterranean algae. Taxon 

36: 755-758. 

ERCEGOVIĆ, A. 1948. Sur quelques algues Phaeophycées 

peu connues ou nouvelles récoltées 

dans le bassin de l’Adriatique moyen (On 

some new or little known brown algae 

collected in the basin of the middle Adriatic.). 

Acta Adriat., 3(5): 33 pp. 

ERCEGOVIĆ, A. 1952. Jadranske cistozire (Sur les 

Cystoseira Adriatiques) (The Adriatic Cystoseira). 

Fauna et Flora Adriatica, 2(1): 1-212. 

ERCEGOVIĆ, A. 1955a. Contribution a la connaissance 

des ectocarpes (Ectocarpus) de 

l’Adriatique moyenne (Contribution to the 

knowledge of Ectocarpus of the middle 

Adriatic). Acta Adriat., 7(5): 74 pp. 

ERCEGOVIĆ, A. 1955b. Contribution a la connaissance 

des pheophycees de l’Adriatique 

moyenne (Contribution to the knowledge of 

the Phaeophyceae of the middle Adriatic). 

Acta Adriat., 7(6): 49 pp. 

ERCEGOVIĆ A. 1957. La flore sous marine de 

l’îlot de Jabuka (Sea flora of Jabuka Pit). 

Acta Adriat., 8: 1-130. 

ERCEGOVIĆ, A. 1966. Pogled na floru i ekologiju 

plitkovodne vegetacije alga u srednjem 

Jadranu (A view on the flora and ecology of 

shallow water algae vegetation in the Middle 

Adriatic). Acta Adriat., 1: 55-75. 

ERCEGOVIĆ, A. 1968. Vegetacija plitkih voda 

(obale) srednjeg Jadrana. Komparativni studij 

plitkovodne i dubinske vegetacije srednjeg 

Jadrana (Shallow waters vegetation in 

the middle Adriatic. Comperative study of 

the shallow and deep waters vegetation in 

the middle Adriatic). Organska produkcija 

morskih ekosistema, Godišnji izvještaj 1967 

(II), IOR Split, 10 pp. + Tabl. 3.(mimeo). 

FLETCHER, R. L., I. M. MUNDA & A. VUKOVIČ. 

1988. Compsonema saxicolium (Kuckuck) 

Kuckuck and Microspongium gelatinosum


Reinke (Scytosiphonaceae, Fucophyceae): 

two new records from the Mediterranean. 

Bot. Mar., 31: 1-8. 

FURNARI, G., M. CORMACI & D. SERIO. 1999. 

Catalogue of the benthic macroalgae of the 

Italian coast of the Adriatic Sea. Bocconea, 

12: 214 pp. 

GALLARDO, T. 1992. Nomenclatural notes on 

some Mediterranean algae, I: Phaeophyceae. 

Taxon, 41: 324-326. 

GIACCONE, G. 1978. Revisione della Flora Marina 

del Mare Adriatico (Revision of the marine 

flora of the Adriatic Sea.). WWF & Ann. 

Parco Mar. Miramare (Trieste), 6: 1-118. 

GERLOFF, J. & M. NIZAMUDDIN. 1975. Bemerkungen 

zur Nomenklatur einiger Arten der 

Gattung Cystoseira C. Ag. (Comments on 

nomenclature of some species of the genus 

Cystoseira C. Ag.) Nova Hedwigia, 26: 341- 

348. 

GUIRY, M.D. & G.M.GUIRY. 2009. AlgaeBase. 

World-wide electronic publication, National 

University of Ireland, Galway. Published on 

line: http://www.algaebase.org; searched on 

29 January 2009. 

HAMEL, G. 1939. Sur la classification des Ectocarpales 

(On the classification of Ectocarpales). 

Botaniska Notiser 1939: 65-70. 

MATJAŠIČ, J., J. ŠTIRN, A. AVČIN, L. KUBIK, T. 

VALENTINČIĆ, F. VELKOVRH & S. VUKOVIČ. 

1975. Flora in favna Severnega Jadrana. Prispevek 

1 (The flora and fauna of the North 

Adriatic. Contribution 1). SAZU, Ljubljana, 

54 pp. 

McNEILL, J., F. R. BARRIE, H. M. BURDET, V. DEMOU- 

LIN, D. J. HAWKSWORTH, K. MARHOLD, D. 

H. NICOLSON, J. PRADO, P. C. SILVA, J. E. 

SKOG, J. H. WIERSEMA & N. J. TURLAND. 2006. 


(Vienna Code) adopted by the Seventh 

International Botanical Congress Vienna, 

Austria, July 2005. A.R.G. Gantner Velag, 

Ruggell, Liechtenstein. 568 pp. 

MUNDA, I. 1954. O rasporeditevi bentonskih alg 

na obrežnem području rta Šila na Krku (The 

distribution of benthic algae in the area of 

Šilo on the Krk Island). Biol. Vestn. Glas. 

Slov. Biol., 3: 78-90. 

MUNDA, I. 1960. On the seasonal distribution 

27 

of benthonic marine algae along the northeastern 

coast of the isle of Krk (Northern 

Adriatic). Nova Hedwigia, 2(1/2): 191-242. 

MUNDA, I. 1979. Some Fucacean associations 

from the vicinity of Rovinj, Istrian coast, 

Northern Adriatic. Nova Hedwigia, 31: 607- 

666. 

PARENTE, M. I., A. I. NETO & R. L. FLETCHER. 2003. 

Morphology and life history of Scytosiphon 

lomentaria (Scytosiphonaceae, Phaeophyceae) 

from the Azores. J. Phycol., 39: 353-359. 

PEDERSEN, P. M. 1978. Culture studies in the pleomorphic 

brown alga Myriotrichia clavaeformis 

(Dictyosiphonales, Myriotrichiaeae). 

Norw. J. Bot., 25: 281-291. 

PIGNATTI, S. & G. GIACCONE. 1967. Flora sommersa 

del Golfo di Trieste (The submerged 

flora in the Gulf of Trieste). Nova Thalassia, 

3 (1): 1-17. 

RIBERA, M. A., A. GÓMEZ-GARRETA, T. GALLAR- 

DO, M. CORMACI, G. FURNARI & G. GIAC- 

CONE. 1992. Check-list of Mediterranean 

Seaweeds. I. Fucophyceae (Warming 1884). 

Bot. Mar., 35: 109-130. 

SOLAZZI, A. 1971. Reperti algologici delle Bocche 

di Cattaro (Algology findings of the 

Kotor Bay.). Thalassia Salentina, 5: 3-18. 

SILVA, P.C., P.W. BASSON & R. L. MOE. 1996. 

Catalogue of the benthic marine algae of 

the Indian Ocean. University of California 

publications in botany, 79: 1259 pp. 

ŠERMAN, D., A. ŠPAN, Z. PAVLETIĆ & B. ANTOLIĆ. 

1981. Phytobenthos of Island Lokrum. Acta 

Bot. Croat., 40: 167-182. 

ŠPAN, A. 1972. Rod Sargassum u Jadranu. Morfološko-sistematska 

i ekološka obrada (The 

genus Sargassum in the Adriatic. Morphological 

and ecological-systematic processing.). 

Ph.D. Thesis. Univerity of Zagreb, 

111 pp. (mimeo) 

ŠPAN, A. 1980. Composition et zonation de la 

flore et végétation benthique de l’ile de 

Hvar (Adriatique Moyenne) (Composition 

and zonation of benthic flora and vegetation 

of the Hvar Island (middle Adriatic).). Acta 

Adriat., 21:169-194. 

ŠPAN, A. 2005. The genus Sargassum in the Adriatic 

Sea: Morphology, systematics and ecology. 

Acta Adriat., 46 (Suppl. 1): 9-80.


ŠPAN, A. & B. ANTOLIĆ. 1981. Fitobentos šireg 

područja Malostonskog zaljeva (Phytebnthos 

of the wide area of Mali Ston Bay). 

Savjetovanje “Malostonski zaljev: prirodna 

podloga i društveno valoriziranje”, 

Dubrovnik, 162-174. 

ŠPAN, A. & B. ANTOLIĆ. 1983. Prilog poznavanju 

fitobentosa Crnogorskog primorja, južni 

Jadran (A contribution to the knowledge of 

phytobenthos of an open region (Crnogorsko 

primorje) in the eastern South Adriatic). 

Stud. Mar. Kot., 13/14: 87-110. 

ŠPAN, A. & B. ANTOLIĆ. 1988. Fitobentos. In: 

Armanda, Z., D. Regner, I. Nožina & Z. 

Vučak (Editors). Ekološka studija nultog 

stanja srednjeg Jadrana za potrebe 

istraživanja podmorja. IOR Split, Studije i 

elaborati, 155: 660-665. Anex: Tabl. A.11.1. 

ŠPAN, A. & B. ANTOLIĆ. 1994. Benthic marine flora 

of Kornati National Park (Kornati Archipelago. 

Middle Adriatic, Croatia). Acta Adriat., 

34:29-44. 

Received: 27 February 2009 

Accepted: 25 February 2010 

ŠPAN, A. & B. ANTOLIĆ. 1997. Sastav, rasprostranjenost 

i stanje bentoske flore na širem 

području Splita (Composition, distribution 

and condition of benthic flora in the Split 

area (middle Adriatic, Croatia)). HAZU, 

Zagreb, Tisuću godina prvoga spomena ribarstva 

u Hrvata, 493-513. 

ŠPAN, A., B. ANTOLIĆ, A. ŠIMUNOVIĆ, I. GRUBELIĆ 

& S. JUKIĆ. 1996. Ecological study of gas 

fields in the northern Adriatic: 12. Ecological 

features of the benthic community. Acta 

Adriat., 37(1/2): 161-194. 

VATOVA, A. 1948. Fenologia delle alghe marine 

di Rovigno (Phenology of marine algae of 

Rovinj.). Nova Thalassia, 1: 43-69. 

VUKOVIČ, A. 1980. Asocijacije morskih bentoških 

alg v Piranskem zalivu (Associations of 

iarine benthic algae in the Piran Bay (Gulf 

of Trieste)). Biol. Vestn. Glas. Slov. Biol., 

28(2): 103-124. 

ZAVODNIK, D., A. ŠPAN, N. ZAVODNIK & B. 

ANTOLIĆ. 1981. Benthos of the western coast 

of the Island Krk (Rijeka Bay, the North 

Adriatic Sea). Thalassia Jugosl., 17(3/4): 

289-340.


Popis morskih bentoskih makroalgi na istočnoj obali Jadrana: 

II. Heterokontophyta: Phaeophyceae 

Boris ANTOLIĆ*, Ante ŠPAN, Ante ŽULJEVIĆ, Vedran NIKOLIĆ, 

Ivana GRUBELIĆ, Marija DESPALATOVIĆ i Ivan CVITKOVIĆ 

Institut za oceanografiju i ribarstvo, Laboratorij za bentos 

Šet. I. Meštrovića 63, 21000 Split, Hrvatska 

* Kontakt adresa, e-mail: antolic@izor.hr 

SAŽeTAK 

U ovom radu iznosimo popis svojti morskih bentoskih makroalgi iz odjeljka smeđih alga (Heterokontophytata: 

Phaeophyceae) koji se temelji na podacima objavljenim između 1948. i 1997. 

godine. Zemljopisno smo istočnu obalu Jadranskog mora podijelili na tri dijela: sjeverni, srednji i 

južni. Navedeno je ukupno 180 vrsta i nižih taksonomskih kategorija smeđih algi među kojima je 

predloženo 12 novih sistematskih kombinacija. Najviše ih je zabilježeno u srednjem (153), manju u 

srednjem (129), a najmanje u južnom (75) Jadranu. 

Ključne riječi: morske bentoske makroalge, Heterokontophyta, Paheophyceae, popis, istočna obala 

Jadrana 

29


ANNEX 

Alphabetical list of algal taxa 

(c = Nomenclutaral change; i = taxon inquirendum; n = Note) 

Acinetospora crinita Cutleria multifida Cystoseira compressa 

Acinetospora pusilla Cylindrocarpus berkeleyi f. rosseta 

Acinetospora species i Cylindrocarpus Cystoseira compressa 

Acinetospora vidovichii microscopicus f. insularum c 

Acrosporangium ralfsioides Cystoseira abrotanifolia Cystoseira compressa 

Ascocyclus magnusii Cystoseira abrotanifolia var. pustulata 

Ascocyclus orbicularis f. insularum c Cystoseira corniculata 

Adriogloia adriatica n5 Cystoseira abrotanifolia var. corniculata 

Agalozonia chilosa n2 subsp. plana Cystoseira corniculata 

Agalozonia melanoidea n1 Cystoseira abrotanifolia f. imperfecta n12 

Agalozonia parvula n3 subsp. pustulata Cystoseira corniculata 

Arthrocladia villosa var. compressa subsp. divergens c 

Asperococcus bullosus Cystoseira adratica Cystoseira corniculata 

Asperococcus compressus subsp. compressa var. divergens c 

Asperococcus echinatus Cystoseira adratica Cystoseira corniculata 

Asperococcus ensiformis var. intermedia subsp. laxior c 

Asperococcus fistulosus Cystoseira adriatica Cystoseira corniculata 

Asperococcus scaber subsp. intermedia var. laxior c 

Asperococcus turneri Cystoseira adriatica Cystoseira crinita 

Castagnea cylindrical subsp. reducta f. crinita 

Castagnea mediterranea Cystoseira adriatica Cystoseira crinita 

Castagnea zosterae f. reducta Cystoseira crinita 

Carpomitra cabrerae Cystoseira adriatica f. semispinosa 

Carpomitra costata subsp. tenuior Cystoseira crinitophylla 

var. costata Cystoseira adriatica Cystoseira discors 

Carpomitra costata f. tenuior Cystoseira discors 

var. dichotoma Cystoseira amentacea f. dubia i 

Choristocarpus tenellus var. spicata Cystoseira discors 

Cladosiphon contortus Cytoseira barbata f. latiramosa 

Cladosiphon cylindricus Cystoseira barbata Cystoseira discors 

Cladosiphon mediterraneus var. barbata subsp. latiramosa 

Cladosiphon zosterae Cystoseira barbata Cystoseira discors 

Cladostephus hirsutus f. insularum f. tenuiramosa 

Cladostephus spongiosum Cystoseira barbata Cystoseira dubia 

f. verticillatum f. punctata i Cystoseira ercegovicii 

Cladostephus verticillatus Cystoseira barbata Cystoseira ercegovicii 

Colpomenia sinuosa var. tophuloidea f. latiramosa 

Compsonema gracile Cystoseira barbata Cystoseira ercegovicii 

Compsonema saxicolum subsp. tophuloidea f. tenuiramosa 

Corynophlaea flaccida Cystoseira barbata Cystoseira erica-marina 

Corynophlaea. umbellata var. tophuloidea Cystoseira fimbriata 

Corynephora umbellata Cystoseira compressa Cystoseira fimbriata 

Cutleria adspersa f. compressa f. rosetta 

Cutleria chillosa Cystoseira compressa Cystoseira foeniculacea 

Cutleria monoica f. plana f. foeniculacea


Cystoseira foeniculacea Dictyota dichotoma Ectocarpus fasciculatus 

f. latiramosa var. intricata var. abbreviatus 

Cystoseira foeniculacea Dictyota dichotoma Ectocarpus flagelliformis 

f. tenuiramosa f. proliferans i Ectocarpus geniculatus 

Cystoseira fucoides Dictyota fasciola Ectocarpus granulosus 

Cystoseira humilis var. fasciola Ectocarpus hauckii 

var. humilis Dictyota fasciola Ectocarpus hinksiae 

Cystoseira humilis var. repens Ectocarpus mitchelliae 

var. myriophylloides Dictyota implexa Ectocarpus irregularis 

Cystoseira. jabukae Dictyota linearis Ectocarpus irregularis 

f. jabukae Dictyota mediterranea subsp. lebeliides c 

Cystoseira. jabukae Dictyota spiralis Ectocarpus lebelii 

f. tenuissima Dilophus fasciola Ectocarpus paradoxoides 

Cystoseira. jabukae Dilophus fasciola Ectocarpus paradoxus 

subsp. tenuissima var. repens Ectocarpus paradoxus 

Cystoseira myriophylloides Dilophus ligulatus var. donatiae c 

Cystoseira opuntioides Dilophus mediterraneus Ectocarpus paradoxus 

Cystoseira pelagosae Dilophus mediterraneus subsp. Lebeliides 

Cystoseira platyramosa var. crassus i Ectocarpus paradoxus 

Cystoseira sauvageauana Dilophus repens f. profonda c 

Cystoseira selagenoides Dilophus spiralis Ectocarpus parvulus 

Cystoseira spinosa Discosporangium Ectocarpus pectinis 

var. compressa mesarthrocarpum Ectocarpus pygmaeus n9 

Cystoseira spinosa Ectocarpus abbreviatus Ectocarpus rufulus 

var. spinosa Ectocarpus adriaticus n7 Ectocarpus sandrianus 

Cystoseira spinosa Ectocarpus adriaticus Ectocarpus siliculosus 

var. tenuior f. maior c var. adriaticus n7 

Cystoseira spinosa Ectocarpus approximatus Ectocarpus siliculosus 

var. squarrosa Ectocarpus arctus n8 var. adriaticus 

Cystoseira spicata Ectocarpus battersii f. maior c 

Cystoseira spicata Ectocarpus battersii Ectocarpus siliculosus 

subsp. crassa i var. mediterraneus var. arctus n8 

Cystoseira spicata Ectocarpus battersiides Ectocarpus siliculosus 

subsp. elegans i Ectocarpus battersiides var. confervoides 

Cystoseira stricta f. maior c Ectocarpus siliculosus 

var. spicata Ectocarpus battersiides var. crassus 

Cystoseira squarrosa f. spongiosessilis c Ectocarpus siliculosus 

Cystoseira zosteroides Ectocarpus battersiides var. dasycarpus 

Dalmatogloia bryozoi n6 f. taoniae c Ectocarpus siliculosus 

Desmarestia adriatica Ectocarpus bombycinus var. divergens i 

Desmarestia ligulata Ectocarpus crinitus Ectocarpus siliculosus 

Dichosporangium repens Ectocarpus confervoides var. elongatus i 

Dictyopteris polypodioides Ectocarpus dalmaticus Ectocarpus siliculosus 

Dictyopteris membranacea Ectocarpus dalmaticus var. megacarpus i 

Dictyota dichotoma f. acinetiformis c Ectocarpus siliculosus 

var. dichotoma Ectocarpus dasycarpus var. pygmaeus n9 

Dictyota dichotoma Ectocarpus fasciculatus Ectocarpus siliculosus 

f. implexa Ectocarpus fasciculatus var. siliculosus 

Dictyota dichotoma var. fasciculatus Ectocarpus siliculosus 

var. implexa var. subulatus n10 

31


Ectocarpus siliculosus Feldmannia lebelii Lithoderma adriaticum 

var. venetus n11 Feldmannia padinae Litosiphon laminariae 

Ectocarpus simpliciusculus Feldmannia paradoxa Leathesia flaccida 

f. simpliciusculus var. paradoxa Leathesia mucosa 

Ectocarpus simpliciusculus Feldmannia paradoxa var. mucosa 

f.reductus (“reducta”) var. donatiae c Leathesia mucosa 

Ectocarpus spinosus Feldmannia paradoxa f. exuberans 

Ectocarpus subulatus n10 f. profonda c Leathesia umbellata 

Ectocarpus valiantei Feldmannia paradoxoides Liebmannia leveillei 

Ectocarpus velutinus n11 Fucus costatus Mesogloia griffithsiana 

Ectocarpus venetus Fucus discors Mesogloia leveillei 

Ectocarpus virescens Fucus visoides Mesogloea leveillei 

Elachista fucicola Giffordia dalmatica Mesogloea vermiculata 

f. fucicola Giffordia geniculata Mesogloia vermiculata 

Elachista fucicola Giffordia granulosa Microspongium 

f. profunda Giffordia haucki gelatinosum n14 

Elachista intermedia Giffordia hinksiae Microsporangium 

var. intermedia Giffordia irregularis kuckuckianum 

Elachista intermedia Giffordia mitchelliae Myriactis adriatica 

var. clavaeformis Giffordia sandriana Myriactis elongata 

Elachista intermedia Giraudia sphacelarioides Myriactis microscopica 

f. profunda Gonodia stellulata Myriactis pulvinata 

Elachista jabukae c Haloglossum Myriactis rigida 

Elachista kuckuckiana compressum Myriactis stelullata 

Elachista neglecta Halopteris filicina Myriactula elongata 

var. neglecta Halopteris scoparia Myriactula microscopica 

Elachista neglecta Halyseris polypodioides Myriactula pulvinata 

var. jabukae c Hecatonema maculans Myriactula rigida 

Elachista neglecta Hecatonema terminale Myriactula rivulariae 

subsp. Jabukae Herponema valiantei Myriactula stellulata 

Elachista stellaris Hincksia dalmatica Myriogloea sciurus i 

Entonema effusum f. dalmatica Myrionema 

Entonema oligosporum Hincksia dalmatica liechtensternii 

Endodictyon infestans f. acinetiformis c Myrionema magnusii 

Feldmannia battersii Hinksia geniculata Myrionema orbiculare 

Feldmannia battersiides Hinksia granulosa Myrionema saxicola 

f. battersiides Hinksia hauckii Myrionema strangulans 

Feldmannia battersiides Hinksia hincksiae Myrionema vulgare 

f. maior c Hinksia mitchelliae Myriotrichia adriatica 

Feldmannia battersiides Hinksia sandriana Myriotrichia 

f. spongiosessilis c Hydroclathrus clathratus clavaeformis 

Feldmannia battersiides Kuckuckia spinosa f. clavaeformis 

f. taoniae c Kuetzingiella battersii Myriotrichia clavaeformis 

Feldmannia caespitula var. battersii f. acycla c 

Feldmannia caespitula Kuetzingiella battersii Myriotrichia 

var. lebelii var. mediterranea protasperococcus 

Feldmannia irregularis Laminaria rodriguezii Myriotrichia repens 

var . irregularis Leptonematella Myriotrichia repens 

Feldmannia irregularis fasciculata f. brevicellularis- i 

var. lebeliides c Leptonema fasciculatum plurilocularis


Myriotrichia repens Sargassum acinarium Sphacelaria cirrosa 

f. brevicellularis- Sargassum hornschuchii var. pennata 

unilocularis i Sargassum linifolium Sphacelaria filicina 

Myriotrichia repens Sargassum salicifolium Sphacelaria furcigera 

f. internodilais- Sargassum salicifolium Sphacellaria fusca 

mixtolocularis i subsp. linifolium n13 Sphacellaria hystrix 

Myriotrichia repens Sargassum vulgare Sphacelaria irregularis 

f. longicellularis- Sargassum vulgare Sphacelaria nana 

unilocularis i f. ercegovicii i Sphacelaria pennata 

Myriotrichia repens Sargassum vulgare Sphacellaria plumula 

f. longicellularis-scoparia i subsp. jabukae i Sphacellaria rigidula 

Myriotrichia repens Sargassum vulgare Sphacelaria scoparia 

f. acycla c var. linifolium Sphacellaria tribuloides 

Nemacystus flexuosus Sargassum vulgare Sporochnus dichotomus 

var. giraudyi subsp. megalophyllum Sporochnus 

Nemacystus ramulosus Sargassum vulgare pedunculatus 

Nereia filiformis var. salicifolium Stictyosiphon adriaticus 

Padina pavonica Sauvageaugloia divaricata Stictyosiphon tortilis 

Padina pavonia Sauvageaugloia Stypocaulon scoparium 

Padinopsis adriatica n4 griffithsiana Stilophora rhizodes 

Petalonia fascia Scytosiphon dotyi Stilophora tenella 

Petalonia zosterifolia Scytosiphon lomentaria Streblonema effusum 

Petrosporangium berkeley Scytosiphon lomentarius Streblonema infestans 

Phaeosphaerium Scytosiphon Streblonema 

liechtensternii simplicissimus oligosporum 

Phyllitis fascia Spatoglossum solieri Streblonema sphaericum 

Phyllitis zosterifolia Spermatochnus Streblonemopsis irritans 

Pilayella litoralis paradoxus Striaria attenuata 

Protasperococcus var. paradoxus Taonia atomaria 

myriotrichiiformis Spermatochnus Zanardinia typus 

Pseudolithoderma paradoxus Zosterocarpus 

adriaticum var. adriaticus oedogonium 

Punctaria latofolia Sphacella subtilissima Zanardinia collaris 

Punctaria tenuissima Sphacelaria britannica Zanardinia prototypus 

Ralfsia verrucosa Sphacelaria cirrosa 

33

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